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Narcinidae by jekky
Subfamily Narcininae Members of this family are commonly known as the numbfishes, proper, and are found almost worldwide in warm temperate and tropical continental and continental insular waters. They are strictly marine and are absent from rivers and lakes. They occur in sandy beaches, muddy enclosed bays, estuaries, off coral reefs and river mouths, and on the upper continental slope to a depth of 1,071 meters. Description Small to medium-sized rays, adult numbfishes range from 15 to 66 cm long, though most are less tha 50 cm. They have oval, rounded, or shovel-shaped pectoral discs and stout tails of equal or longer length. The snout is moderately elongated, rounded or rounded-angular, differing from the narkids in being supported by broad rostral cartilages. The mouth is straight, with stout, elongated, and highly protrusible jaws and a prominent groove around its periphery. The nostrils are just forward of the mouth and connected to it by a broad groove; the nasal flaps are short but merged into a broad nasal curtain that overlaps the mouth. The teeth are small have a single moderate cusp; the teeth and tooth bands are exposed when the mouth is closed (except in Diplobatis). There are two prominent dorsal fins of roughly equal size, and a large caudal fin. Numbfishes vary in color from whitish, yellowish, brownish, grey-brown, greenish, reddish, or black above, either plain or with small to large spots, blotches, bars or lines, sometimes forming complex eye-shaped spots or ocelli on the pectoral fins. They are usually white underneath, or black in deep-water species. The large, kidney-shaped electric organs are at the base of the pectoral fins and visible through the skin. Biology and ecology Numbfishes are slow-swimming bottom-dwellers that feed on small fishes and invertebrates off the bottom; their protrusible jaws aid in removing prey from the substrate. They can generate a moderate shock if disturbed and contact is made with the electric organs; the electrical discharges of narcinids have been measured at 8-37 volts, much less than the electric rays of the genus Torpedo. All species are ovoviviparous, with eggs hatching inside the mother. Species Genus Benthobatis Benthobatis kreffti Rincon, Stehmann & Vooren, 2001 Blind torpedo, Benthobatis marcida Bean & Weed, 1909 Dark blind ray, Benthobatis moresbyi Alcock, 1898 Genus Diplobatis Diplobatis colombiensis Fechhelm & McEachran, 1984 Diplobatis guamachensis Martn Salazar, 1957 Ocellated electric ray, Diplobatis ommata (Jordan & Gilbert, 1890) Painted electric ray, Diplobatis pictus Palmer, 1950 Genus Discopyge Apron ray, Discopyge tschudii Heckel, 1846 Genus Narcine Narcine atzi Carvalho & Randall, 2003 Lesser electric ray, Narcine bancroftii (Griffith & Smith, 1834) Brazilian electric ray, Narcine brasiliensis (Olfers, 1831) Shortlip electric ray, Narcine brevilabiata Bessednov, 1966 Brown numbfish, Narcine brunnea Annandale, 1909 Giant electric ray, Narcine entemedor Jordan & Starks, 1895 Largespotted numbfish, Narcine indica Henle, 1834 Narcine insolita Carvalho, Sret & Compagno, 2002 Narcine lasti Carvalho & Sret, 2002 Narcine leoparda Carvalho, 2001 Chinese numbfish, Narcine lingula Richardson, 1846 Darkfinned numbfish, Narcine maculata (Shaw, 1804) Narcine oculifera Carvalho, Compagno & Mee, 2002 Tonkin numbfish, Narcine prodorsalis Bessednov, 1966 Slender electric ray, Narcine rierai (Lloris & Rucabado, 1991) Tasmanian numbfish, Narcine tasmaniensis Richardson, 1841 Blackspotted numbfish, Narcine timlei (Bloch & Schneider, 1801) Vermiculate electric ray, Narcine vermiculatus Breder, 1928 Banded numbfish, Narcine westraliensis McKay, 1966 Subfamily Narkinae Members of the subfamily Narcininae are commonly known as sleeper rays. They are restricted to the temperate and tropical Indo-West Pacific from South Africa to Japan to Indonesia, and are exclusively marine and are absent from freshwater habitats. They occur from the intertidal zone to the continental shelf and the upper continental slope to a depth of 350 meters, favoring soft-bottomed habitats. Description Adult sleeper rays range in size from 8 to 46 cm in length and have flattened oval, circular, or pear-shaped pectoral fin discs. They have naked skin, without dermal denticles or thorns. The snout is moderately elongate and broadly rounded, with the rostal cartilage reduced to a slender medial rod. This distinguishes the narkids from the family Narcinidae, which have somewhat longer snouts supported by broad rostral cartilages. The mouth is straight, very narrow, and not highly protrusible, with strong labial folds and a weak groove around the periphery. The nostrils are placed just forward of the mouth and are connected to it by a broad nasoral groove; the nasal flaps are long and fused into a central nasal curtain that overlaps the mouth. There are a pair of large kidney-shaped electric organs at the base of the pectoral fins, visible through the skin. The genera Heteronarce and Electrolux have two large dorsal fins, Narke, Crassinarke, and Typhlonarke have a single dorsal fin, and Temera has none. No other group of electric rays have genera with a variable number of dorsal fins. The tail is moderately long, with a large, rounded, symmetrical caudal fin. The color is usually brown or reddish brown above, white or brownish below. There may be spots, blotches, or bands on the dorsal surface, but most species lack complex color patterns or ocelli. Biology and ecology Sleeper rays are bottom-dwelling, slow-moving fishes often found on soft mud or sand. From those species that have been examined, their diet may be restricted to small, soft-bodied, bottom invertebrates, such as polychaete worms. They can generate a strong electric shock as a defense mechanism. Reproduction is ovoviviparous, with eggs hatching inside the mother. Species Genus Crassinarke Sleeper torpedo, Crassinarke dormitor Takagi, 1951 Genus Electrolux Ornate sleeper ray, Electrolux addisoni Compagno & Heemstra, 2007 Genus Heteronarce Elat electric ray, Heteronarce bentuviai (Baranes & Randall, 1989) Natal electric ray, Heteronarce garmani Regan, 1921 Soft electric ray, Heteronarce mollis (Lloyd, 1907) Quilon electric ray, Heteronarce prabhui Talwar, 1981 Genus Narke Cape numbfish, or Onefin Electric ray, Narke capensis (Gmelin, 1789) Spottail sleeper ray, Narke dipterygia (Bloch & Schneider, 1801) Japanese sleeper ray, Narke japonica (Temminck & Schlegel, 1850) Genus Temera Finless sleeper ray, Temera hardwickii Gray, 1831 Genus Typhlonarke Blind electric ray, Typhlonarke aysoni (Hamilton, 1902) Oval electric ray, Typhlonarke tarakea Phillipps, 1929 References ^ Narcinidae (TSN 564006). Integrated Taxonomic Information System. Retrieved on March 20 2006. ^ a b Martin, R. Aidan. Electric Rays. ReefQuest Centre for Shark Research. Retrieved on October 12, 2008. ^ Nelson, J.S. (2006). Fishes of the World (fourth ed.). John Wiley. pp. 69-82. ISBN 0471250317. ^ a b c d Compagno, L.J.V. and Last, P.R. (1999). Narcinidae. Numbfishes. p. 1433-1437. In: K.E. Carpenter and V.H. Niem (eds.) FAO identification guide for fishery purposes. The living marine resources of the Western Central Pacific. Rome: Food and Agricultural Organization. ^ a b Hamlett, William C. (1999). Sharks, Skates, and Rays: The Biology of Elasmobranch Fishes. Baltimore and London: JHU Press. ISBN 0-8018604-8-2. ^ a b c Compagno, L.J.V. and Last, P.R. (1999). Narkidae. Sleeper rays. p. 1443-1446. In: K.E. Carpenter and V.H. Niem (eds.) FAO identification guide for fishery purposes. The living marine resources of the Western Central Pacific. Rome: Food and Agricultural Organization. Categories: Torpediniformes
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